ALAN B. FRANKLIN, Ph.D.
Colorado Cooperative Fish &Wildlife Research Unit
201 Wagar Building
Colorado State University
Fort Collins, CO 80523
970-491-4870 (voice)      970-491-1413 (FAX)
alanf@cnr.colostate.edu

EDUCATION

  • Ph.D., Wildlife Biology (1997), Colorado State University, Fort Collins, Colorado.
    Dissertation:    Factors affecting temporal and spatial variation in northern spotted owl populations
    Major Advisor: Dr. David R. Anderson
  • M.S., Natural Resources (1987), Humboldt State University, Arcata, California.
    Thesis:                 Breeding biology of great gray owls in southeastern     Idaho and western Wyoming
    Major Advisor:    Dr. R. J. Gutiérrez
  • B.S., Wildlife Science (1978), Cornell University, Ithaca, New York.

POSITIONS HELD

Professional Employment:

  • Research Associate (1998 - present), Colorado Cooperative Fish and Wildlife Research Unit, Colorado State University, Fort Collins, Colorado. Research appointment working on a variety of projects.
  • Project Leader (1985 - present), Humboldt State University Foundation, Humboldt State Univ., Arcata, California.  Full-time position directing research on the population ecology of northern spotted owls.   Responsible for all phases of administration and publication of results.
  • Consultant (1983 - present).  Provided professional consulting (paid and unpaid) to government agencies, non-profit organizations and private firms on a variety of topics.
  • Research Associate (1983 - 1984), Humboldt State University Foundation, Humboldt State Univ., Arcata, California. Full-time research position participating in research on dispersal of juvenile northern spotted owls.
  • President (1981 - 1984), Wildlife Research Institute, Inc., Victor, Idaho. Founded and directed non-profit, tax-exempt organization which promoted and conducted wildlife research.
  • Wildlife Technician (1980), Targhee National Forest, Ashton, Idaho. Assisted in developing District-wide wildlife habitat map.
  • Technical Aide (1979), Library of Natural Sounds, Laboratory of Ornithology, Cornell University, Ithaca, New York. Cataloged and assisted in analysis of world-wide collection of avian vocalizations.
  • Research Assistant (1972 - 1974), Charles Darwin Research Station, Galapagos Islands, Ecuador. Assisted with field projects including ecology of Darwins finches and seabirds, marine mammals, and volcanology. Routine duties included mist-netting and banding birds, operating weather and seismic stations, and curating museum collections.

Professional Affiliations:

  • Adjunct Assistant Professor (2000 - present), Department of Fisheries and Wildlife, University of Minnesota, St. Paul, Minnesota.
  • Adjunct Professor (1998 - present) Department of Wildlife, Humboldt State University, Arcata, California. 
  • Faculty Associate (1987 - 1998), Department of Wildlife, Humboldt State University, Arcata, California.

RESEARCH EXPERIENCE

  • Population Ecology of Northern Spotted Owls (1985 - present), Humboldt State University Foundation, Arcata, California. Involved in long-term study on population dynamics of northern spotted owls in northwestern California.  Developed study design, data collection and data analysis procedures.
  • Long-term Trends in Survival in North American Birds (1997- present), Colorado Cooperative Fish and Wildlife Research Unit, Fort Collins, Colorado. Involved in analysis of >25-year band recovery data for avian species in North America to determine long-term trends in survival.
  • Modeling Golden Eagle Populations (1994 - 1997). National Renewable Energy Laboratory, Golden Colorado. Worked with scientific team to develop data-based population model to assess effects of commercial wind turbines on population trends in golden eagles at Altamont Pass, California
  • Statistical Design of Monitoring Programs for Spotted Owls (1992 - present). National Park Service, U. S. Fish and Wildlife Service, and U. S. Forest Service. Involved with developing statistical designs to monitor northern and Mexican spotted owl populations using capture-recapture and quadrat sampling designs.
  • Meta-analysis and Population Monitoring (1992 - 1993). Colorado Cooperative Fish and Wildlife Research Unit, Colorado State University, Fort Collins, Colorado. Explored use of meta-analysis as a tool for incorporating the results of research studies in wildlife monitoring programs.
  • Dispersal of Juvenile Northern Spotted Owls (1983 - 1985), Humboldt State University Foundation, Arcata, California. Participated in radio-telemetry study to measure dispersal patterns in juvenile northern spotted owls.
  • Breeding Biology of Great Gray Owls (1980 - 1982), Dept. Wildlife, Humboldt State University, Arcata, California.   Examined breeding distribution, nesting behavior, and breeding biology of great gray populations in Idaho and Wyoming.
  • Distribution of Furbearer Populations (winter 1979), Bridger-Teton National Forest, Jackson, Wyoming. Assessed past and present distributions of endangered or extinct populations of lynx, fisher, wolverine and wolves in southeast Idaho.
  • Ecology of the Galapagos Rail, (1974) Galapagos Islands, Ecuador. Conducted research on natural history and behavior of Galapagos rails on Santa Cruz Island.

TEACHING EXPERIENCE

Courses Taught:

  • WLDM 485: Management of Endangered Species (Spring 1991; 1 credit). Department of Wildlife, Humboldt State University, Arcata, California.
  • WLDM 323: Ecology of Raptors (Fall 1990; 3 credits). Department of Wildlife, Humboldt State University, Arcata, California
  • CE 859: Natural History of the Spotted Owl (Spring 1986: 1 credit). Chico State University Extension, Chico, California. .
  • UCE 430.6: Natural History of the Spotted Owl (Spring 1986: 1 credit), University of California Extension, Santa Cruz, California.
  • WLDM 199: Research Techniques for Northern Spotted Owls (Spring 1985, Spring 1986: 2 credits), Department of Wildlife, Humboldt State University, Arcata, California.

Workshop Instructor:

  • Intermediate-level Workshop on Program MARK (1999: 5 days). Colorado Cooperative Fish and Wildlife Research Unit, Fort Collins, Colorado.
  • Endangered Species Act Recovery Planning and Implementation (1996: 3 days). U.S. Fish & Wildlife Service National Education Training Center, San Antonio, Texas.
  • International Workshop on Design and Analysis of Distance Sampling Data - Theory and Application (1996: 7 days). University of Natal, Pietermaritzburg, South Africa.
  • Wildlife Management Short Course (1996: 1 day). Colorado State University, Fort Collins, Colorado.
  • International Workshop: Design and Analysis of Distance Sampling Data - Theory and Application (1993: 3 days). Colorado Cooperative Fish and Wildlife Research Unit, Fort Collins, Colorado.
  • Methods for Monitoring and Inventorying Spotted Owls (Spring 1987, 1988: 5 days), Natural Resources Institute, Humboldt State University, Arcata, California.

Workshop Organizer:

  • Workshop on Analysis of Demographic Rates of California Spotted Owls (2001: 5 days). U. S. Forest Service, Colorado State University, Fort Collins, CO. Organized multi-agency workshop to analyze region-wide California spotted owl demography data conducted by a number of different researchers.
  • Estimating Population Parameters from Capture-Recapture Data using Program MARK (2001: 5 days). Department of Wildlife, Humboldt State University, Arcata, California.  Organized and co-taught (with Gary White and Steven Dinsmore of Colorado State University) intensive workshop on using mark-recapture estimators in wildlife population studies.
  • Introduction to Mark-recapture and Program MARK (2000; 1 day). Department of Wildlife, Humboldt State University, Arcata, California. Organizer.
  • Program MARK: Techniques for Monitoring Wildlife Populations (1999; 1 day). Second International Wildlife Congress, Budapest, Hungary. Co-organizer.
  • Workshop on Analysis of Demographic Rates of Northern Spotted Owls (1998: 9 days). U. S. Geological Survey, Biological Resources Division, Corvallis, Oregon. Co-organizer.

Guest Lectures:

  • WLDM 478: Wildlife Populations. Humboldt State University, Arcata, California
  • WLDM 300: Wildlife Ecology. Humboldt State University, Arcata, California
  • WLDM 110: Introduction to Wildlife. Humboldt State University, Arcata, California
  • FW100: Wildlife Biology. Colorado State University, Fort Collins, Colorado
  • FW 555: Conservation Biology.  Colorado State University, Fort Collins, Colorado
  • FW 469: Mgmt of Large Mammals. Colorado State University, Fort Collins, Colorado
  • NR 320: Natural Resource Policy. Colorado State University, Fort Collins, Colorado

Teaching Assistantships:

  • BIOL 475: Ornithology (Fall 1978: 4 credits), Department of Ecology and Systematics, Cornell University, Ithaca, New York. (under T. Cade).

Graduate Students:

            Major Advisor to:

  • Stephanie Waldo, M.S. candidate, Department Wildlife, Humboldt State University, Arcata, California. Discrimination of neighbors and strangers by territorial northern spotted owls.
  • Jennifer Van Gelder, M.S. candidate, Department Wildlife, Humboldt State University, Arcata, California. Geographic variation in spotted owl vocalizations.

            Committee Member to:

  • Armand Gonzales, M.S. candidate, Department Wildlife, Humboldt State University, Arcata, California. Comparison of actual versus predicted habitat use by the northern spotted owl in California using a geographic information system.
  • Tih-Fen Ting, M.S. degree (1998) Department Wildlife, Humboldt State University, Arcata, California. The thermal environment of northern spotted owls in northwestern California: possible explanations for use of interior old growth and coastal early successional stage forest.

HONORS AND AWARDS

  • Outstanding Publication in Wildlife Ecology and Management (Monograph Category), The Wildlife Society, Bethesda, Maryland.  Received  for “excellence in scientific writing characterized by originality of research or thought and a high scholastic standard in the manner of presentation” for Franklin et al. 2000 Ecological Monographs 70:539-590.
  • Elective Member, American Ornithologists’ Union, Washington, DC. (2000) Election in this membership class was bestowed in recognition of my contributions to ornithology.
  • Conservation Service Award, U. S. Department of the Interior, Washington, DC. (1999) Received award and citation as a member of the Mexican Spotted Owl Recovery Team. Highest honor given to civilians by the Department of Interior.
  • Elected Member, Honor Society of Phi Kappa Phi (1998)
  • Monograph of the Year Award (1997), The Wildlife Society, Bethesda, Maryland. Authored four papers included in Studies in Avian Biology 17 that received the award.
  • Travel Award (1995), Department of Fishery and Wildlife and College of Natural Resources, Colorado State University, Fort Collins, Colorado. Received funds for travel to South Africa.
  • Certificate of Appreciation (1994), Six Rivers National Forest, Willow Creek, California. For integrating research with management of spotted owls.
  • Certificate of Appreciation (1993), U. S. Forest Service, Washington, D.C. For technical work on the Report of the Scientific Analysis Team: Viability Assessments and Management Considerations for Species Associated with Late-Successional and Old-growth Forests of the Pacific Northwest.
  • Certificate of Appreciation (1993), U. S. Forest Service, Washington, D.C. In recognition for service on the President's Ecosystem Management Assessment Team.
  • Certificate of Appreciation (1990), Interagency Spotted Owl Scientific Committee, Portland, Oregon. For assistance in preparation of a conservation strategy for the northern spotted owl.
  • Research Award (1981 and 1982), Sigma Xi Scientific Research Society, New Haven Connecticut. Received two awards for thesis research.

CONSULTING

Federal & State Agencies:

  • Preble’s Meadow Jumping Mouse Recovery Team (2000), Colorado Division of Wildlife, Fort Collins, Colorado. Formally consulted with Team members on conservation biology issues and reserve design strategies to meet recovery goals.
  • U. S. Forest Service Sierra Nevada Framework Team (2000), Sacramento, California.  Consulted with members of the team about designing a monitoring strategy for California spotted owls.
  • Oregon Cooperative Fish & Wildlife Research Unit (1999), Oregon State University, Corvallis, Oregon.  Consulted with biologists on designing and implementing study to examine the effects of landscape habitat characteristics on northern spotted owl demography.
  • U. S. Forest Service (1999), Sierra Nevada Framework Science Team, Sacramento, California.  Assisted with evaluating environmental impact statement alternatives and demographic modeling approaches for the California spotted owl.
  • U. S. Fish and Wildlife Service (1998-1999), Arcata, California. Consulted on developing habitat map for northern spotted owls across their range in California, using model selection procedures and data on occupancy
  • California Department of Fish and Game (1997), Eureka, California. Reviewed proposed monitoring plan for northern spotted owls on Pacific Lumber Company lands.
  • U. S. Department of Energy (1994 - 95). National Renewable Energy Laboratory, Golden, Colorado. Served as paid consultant to design study for assessing impacts of wind turbine farms on golden eagle populations.
  • U. S. Forest Service (1993). Rocky Mountain Forest and Range Experiment Station, Flagstaff, Arizona. Estimated survival rates for Mexican spotted owls from mark-recapture and telemetry data.
  • U. S. Forest Service (1993) Six Rivers National Forest, Eureka, California. Participated in developing a proposed adaptive ecosystem management project for managing wildlife habitat.
  • U. S. Forest Service (1993 and 1998). Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado and Oregon State University, Corvalis, Oregon. Assisted in implementation of two workshops to analyze region-wide northern spotted owl demography data.
  • National Park Service (1992), Olympic National Park, Port Angeles, Washington. Served as paid committee member to develop study design for estimating density of northern spotted owls on National Park lands.
  • U. S. Forest Service (1992), Rocky Mountain Forest and Range Experiment Station, Flagstaff, Arizona. Assisted in implementation of Mexican spotted owl radio-telemetry project in New Mexico.
  • U. S. Forest Service (1992), Southwest Region, Albuquerque, New Mexico. Served as paid reviewer for proposed management strategy for Mexican Spotted Owls in Region 3 of the U.S. Forest Service.
  • U. S. Fish and Wildlife Service (1988 - 1991), Portland, Oregon. Consulted with members of Status Review Team, Recovery Team and management staff concerning northern spotted owl biology and management.

Non-Profit Organizations:

  • Sustainable Ecosystem Institute (1999), Portland, Oregon.  Assisted with peer-review program for federal natural resource decisions by recruiting potential reviewers and serving as a peer-reviewer.
  • Sierra Club (1998). Oakland, California. Consulted on review of northern spotted owl provisions of proposed Pacific Lumber Company Habitat Conservation Plan
  • Film Productions (1988 - 1991), British Broadcasting Co., American Broadcasting Co., Nature Film Co., Green TV Productions. Consulted in production of news and full-length film productions on old-growth ecosystems such as Good Morning America, "Land of the Eagle", "Life Force" and "Heart of the Wood".
  • The Wilderness Society (1987), Washington, DC. Assessed proposed management of spotted owls in the Shasta-Trinity National Forest Plan (with R. J. Gutiérrez).

PROFESSIONAL ACTIVITIES

Professional Societies:

  • Memberships: The Wildlife Society, Ecological Society of America, Society of Conservation Biologists, American Ornithologists Union, Cooper Ornithological Society, Raptor Research Foundation
  • Board Member (2001), Biometrics Working Group, The Wildlife Society, Bethesda, Maryland.  Served as a member of the Board of Directors and as the Audit, Elections and Nominating Committee Chair.
  • Symposium Organizer (1997), 4th Annual Conference of the Wildlife Society, Snowmass, Colorado. Organized (with T. M. Shenk) all-day symposium entitled 'How to Practice Safe Modeling: the Interpretation and Application of Models in Resource Management'.
  • Session Organizer (2000) EURING 2000 Conference, Marshall, California.  Organized half-day session (with Rhys Green) on “Direct estimation of Lambda from Capture-Recapture Studies”.
  • Session Moderator (1999 - 2001) Moderated Conservation and Management session at 1999 Raptor Research Foundation Annual Meeting, La Paz, Mexico and Wildlife Population Dynamics, Estimation, and Modeling session at The Wildlife Society 8th Annual Conference, Reno, Nevada.
  • Reviewer, Scientific Journals (1988 - present). Acted as peer-reviewer for papers submitted to Journal of Wildlife Management, Wildlife Monographs, Ecology, Ecological Applications, Biometrics, Wildlife Biology, The Auk, The Condor, North American Bird Bander, Journal of Raptor Research, Journal of Applied Statistics, Northwestern Naturalist, Conservation Biology, Forest Science, Wilson Bulletin, Journal of Agricultural, Biological and Environmental Statistics and 2 books.

Conservation Planning:

  • Member, Scientific Review Panel (2000 - present), Sustainable Ecosystem Institute, Portland, Oregon. Serve on scientific review panel to provide oversight on Pacific Lumber Company in compliance of their habitat conservation plan to monitor marbled murrelet and northern spotted owl populations.
  • Member, Northern Spotted Owl Effectiveness Monitoring Team (1998 - present). U. S. Forest Service, Portland, Oregon. Member of scientific team to implement range-wide monitoring program for northern spotted owls.
  • Member, Mexican Spotted Owl Recovery Team (1993 - 2001), U. S. Fish and Wildlife Service, Albuquerque, New Mexico. Designated member of recovery team for a federally-listed threatened species.
  • Member, Northern Spotted Owl Viability Panel (1993), Interagency Forest Ecosystem Management Team, Portland, Oregon. Assessed alternative options for ecosystem management plans with respect to northern spotted owl populations
  • Member, Interagency Spotted Owl RD&A Technical Group (1986 - 1990), U. S. Forest Service, Portland, Oregon. Participated in annual meetings to guide and standardize spotted owl research and population monitoring.
  • Expert Testimony, Interagency Spotted Owl Scientific Committee (1989), U.S. Forest Service, Portland, Oregon. Presented results of spotted owl research to scientific panel.
  • Reviewer, Spotted Owl Management Plans (1981 - 1992) Provided input and reviews of spotted owl management plans for the U.S. Forest Service, Simpson Timber Company, Pacific Lumber Company, and the Hoopa Tribal Council.
  • Expert Testimony, Spotted Owl Advisory Panel (1985), American Ornithologists Union, Sacramento, California. Presented oral and written testimony on spotted owl research.

Non-Profit Organizations:

  • Expert Witness, (1989), Sierra Club Legal Defense Fund, Seattle, Washington. Provided affidavits, depositions and courtroom testimony in litigation with the Bureau of Land Management.
  • Expert Witness, (1989), Sierra Club Legal Defense Fund, San Francisco, California. Provided affidavits in litigation with the U.S. Forest Service.
  • Expert Witness, (1988), Environmental Protection Information Center, Garberville, California. Provided affidavits in litigation with the U.S. Forest Service.
  • Member, Board of Directors, (1987 - 1988), Redwood Region Audubon Society, Arcata, California.

PUBLICATIONS

Books:

  • Shenk, T. M. and A. B. FRANKLIN, editors. 2001. Modeling in natural resource management: development, interpretation, and application. Island Press, Washington, DC.

Scientific Papers:

  • Bond, M. L., R.. J. Gutiérrez, A. B. FRANKLIN, W. S. LaHaye, C. A. May, and M. E. Seamans. 2003 Survival and movements of spotted owls immediately after wildfire. Wildlife Society Bulletin. In Press.
  • Parmenter, R. R., T. L. Yates, D. R. Anderson, K. P. Burnham, J. L. Dunham, A. B. FRANKLIN, M. T. Friggens, B. Lubow, M. Miller, G. S. Olson, C. A. Parmenter, J. Pollard, E. Rexstad, T. M. Shenk, T. R. Stanley, and G. C. White. 2003. Small mammal density estimation: a field comparison of grid-based versus web-based density estimators using enclosed rodent populations. Ecology. In Press.
  • FRANKLIN, A. B., B. R. Noon, and T. L. George. 2002. What is habitat fragmentation? Studies in Avian Biology 25:20-29.
  • FRANKLIN, A. B. and R. J. Gutiérrez. 2002. Spotted owls, forest fragmentation, and forest heterogeneity. Studies in Avian Biology. 25:204-221.
  • FRANKLIN, A. B., D. R. Anderson, and K. P. Burnham. 2002. Estimation of long-term trends and variation in avian survival probabilities using random effects models. Journal of Applied Statistics 29:267-287. (reprint)
  • Noon, B. R. and A. B. FRANKLIN. 2002. Scientific research and the spotted owl: opportunities for major contributions to avian population ecology. Auk 119:311-320. (reprint)
  • White, G. C., A. B. FRANKLIN, and T. M. Shenk. 2002. Estimating parameters of PVA models from data on marked animals. Pages 169-190 In S. R. Beissinger and D. R. McCullough, editors. Population Viability analysis. Population Viability Analysis. University of Chicago Press, Chicago, Illinois. (reprint)
  • Block, W. M., A. B. FRANKLIN, J. P. Ward, Jr., J. L. Ganey, and G. C. White. 2001. Design and implementation of monitoring studies to evaluate the success of ecological restoration on wildlife. Restoration Ecology 9:293-303. (reprint)
  • Franklin, A. B. 2001. Exploring ecological relationships in survival and estimating rates of population change using program MARK. Pages 290-296 In R. Field, R. J. Warren, H. Okarma, and P. R. Sievert, editors.  Wildlife, land, and people: priorities for the 21st century. Proceedings of the Second International Wildlife Management Congress. The Wildlife Society, Bethesda, Maryland. (reprint)
  • FRANKLIN, A. B., T. M. Shenk, D. R. Anderson, and K. P. Burnham. 2001. Statistical model selection: the alternative to null hypothesis testing. Pages 75-90 In: T. M. Shenk and A. B. Franklin, editors. Modeling in natural resource management: development, interpretation, and application. Island Press, Washington, D.C.
  • Shenk, T. M., and A. B. FRANKLIN. 2001. Models in natural resource management: an introduction. Pages 1-8 in T. M. Shenk and A. B. Franklin, editors. Modeling in natural resource management: development, interpretation, and application. Island Press, Washington, D.C.
  • FRANKLIN, A. B., D. R. Anderson, R. J. Gutiérrez, and K. P. Burnham. 2000. Climate, habitat quality, and fitness in northern spotted owl populations in northwest California. Ecological Monographs. 70:539-590. (reprint)
  • Anderson, D. R., K. P. Burnham, A. B. FRANKLIN, R. J. Gutiérrez, E. D. Forsman, R. J. Anthony, G. C. White, and T. M. Shenk. 1999. A protocol for conflict resolution in analyzing empirical data related to natural resource controversies. Wildlife Society Bulletin 27:1050-1058.
  • White, G. C., W. M. Block, J. P. Ward, Jr., J. L. Ganey, A. B. FRANKLIN, S. L. Spangle, R. J. Vahle, S. E. Rinkevich, W. H. Moir, F. P. Howe, and J. L. Dick, Jr. 1999. Science versus political reality in delisting criteria for a threatened species: the Mexican spotted owl experience. Transactions of the North American Wildlife and Natural Resources Conference 64:292-306.
  • Carlson, P. C., W. S. Lahaye, and A. B. FRANKLIN. 1998. Incestuous behavior in spotted owls. Wilson Bulletin 110:562-564
  • FRANKLIN, A. B., R. J. Gutiérrez, B. R. Noon, and J. P. Ward, Jr. 1996. Demographic characteristics and trends of northern spotted owls in northwestern California. Studies in Avian Biology 17:83-91.
  • FRANKLIN, A. B., D. R. Anderson, E. D. Forsman, K. P. Burnham, and F. F. Wagner. 1996. Methods for collecting and analyzing demographic data on the northern spotted owl. Studies in Avian Biology 17:12-20.
  • Kristan, D. M., R. J. Gutiérrez, and A. B. FRANKLIN. 1996. Adaptive significance of growth patterns in juvenile spotted owls. Canadian Journal of Zoology 74:1882-1886.
  • Raphael, M. G., R. G. Anthony, S. DeStefano, E. D. Forsman, A. B. FRANKLIN, R. Holthausen, E. C. Meslow, and B. R. Noon. 1996. Use, interpretation, and implications of demographic analyses of northern spotted owl populations. Studies in Avian Biology 17:102-112.
  • Gutiérrez, R. J., E. D. Forsman, A. B. FRANKLIN, and E. C. Meslow. 1996. History of demographic studies in the management of spotted owls. Studies in Avian Biology 17:6-11.
  • Forsman, E. D., A. B. FRANKLIN, F. M. Oliver, and J. P. Ward. 1996. A color band for spotted owls. Journal of Field Ornithology 67:507-510.
  • Shenk, T. M., A. B. FRANKLIN, and K. R. Wilson. 1996. A model to estimate the annual rate of golden eagle population change at the Altamont Pass Wind Resource Area, Pages 47-56 in LGL Environmental Research Associates, editors. Proceedings of the National Avian - Wind Power Planning Meeting II, Washington D.C.
  • FRANKLIN, A.B. and T.M. Shenk. 1995. Meta-analysis as a tool for monitoring wildlife populations. Pages 484-487 In J. A. Bissonette and P. R. Krausman, editors. Integrating people and wildlife for a sustainable future. Proceedings of the International Wildlife Management Congress. The Wildlife Society, Bethesda, Md.
  • Gutiérrez, R. J., A. B. FRANKLIN, and W. S. LaHaye. 1995. The spotted owl. The Birds of North America No. 179. 28pp.
  • Hunter, J. E., R. J. Gutiérrez, and A. B. FRANKLIN. 1995. Habitat configuration around spotted owl sites in northwestern California. Condor 97:684-693.
  • Ward, J. P., Jr, A. B. FRANKLIN, S. E. Rinkevich, and F. Clemente. 1995. Distribution and abundance of Mexican spotted owls. Chapter 1 (14pp.) in U. S. Fish and Wildlife Service. Mexican Spotted Owl Recovery Plan, Volume II. Albuquerque, New Mexico.
  • White, G. C., A. B. FRANKLIN, and J. P. Ward, Jr. 1995. Population biology. Chapter 2 (25pp.) in U. S. Fish and Wildlife Service. Mexican Spotted Owl Recovery Plan, Volume II. Albuquerque, New Mexico.
  • Keitt, T., A. B. FRANKLIN, and D. Urban. 1995. Landscape analysis and metapopulation structure. Chapter 3 (16pp.) in U. S. Fish and Wildlife Service. Mexican Spotted Owl Recovery Plan, Volume II. Albuquerque, New Mexico.
  • Hunter, J. E., R. J. Gutiérrez, A. B. FRANKLIN, and D. Olson. 1994. Ectoparasites of the spotted owl. Journal of Raptor Research 28:232-235.
  • Young, K.E., A.B. FRANKLIN, and J.P. Ward. 1993. Infestation of northern spotted owls by hippoboscid flies. Journal of Wildlife Diseases 29(2): 278-283.
  • FRANKLIN, A.B. 1992. Population regulation in northern spotted owls: theoretical implications for management. Pages 815-827 In D. R. McCullough and R. J. Barrett, editors. Wildlife 2001: Populations. Elsevier Science Publications, London, England.
  • Blakesley, J. A., A.B. FRANKLIN and R. J. Gutiérrez. 1992. Spotted owl roost and nest site selection in northwest California. Journal of Wildlife Management 56:388-392.
  • Moen, C.A., A.B. FRANKLIN and R. J. Gutiérrez. 1991. Age determination of subadult northern spotted owls in northwest California. Wildlife Society Bulletin 19:489-493.
  • Ward, J.P., A.B. FRANKLIN and R. J. Gutiérrez. 1991. Estimating spotted owl abundance using search time and regression. Ecological Applications 1:207-214.
  • FRANKLIN, A.B., J.P. Ward, R. J. Gutiérrez and G.I. Gould. 1990. Density of northern spotted owls in northwest California. Journal of Wildlife Management 54:1-10.
  • Blakesley, J. A., A.B. FRANKLIN and R. J., Gutiérrez. 1990. Sexual dimorphism in northern spotted owls from northwest California. Journal of Field Ornithology 61:95-112.
  • FRANKLIN, A.B. 1988. Breeding biology of the Great Gray Owl in southeastern Idaho and northwestern Wyoming. Condor 90: 689-696.
  • Gutiérrez, R. J., A.B. FRANKLIN, B. LaHaye, V. Meretsky and P. Ward. 1985. Juvenile Spotted Owl dispersal in northwestern California: preliminary results. Pages 60-65 In R. J. Gutiérrez and A. B. Carey, editors. Ecology and management of Spotted Owls in the Pacific Northwest. U. S. Forest Service General Technical Report PNW-185.
  • FRANKLIN, A.B., D. Clark and D. Clark. 1979. Ecology and behavior of the Galapagos Rail. Wilson Bulletin 91: 202-221.

Book Reviews:

  • FRANKLIN, A. B. 1995. The wisdom of the spotted owl: policy lessons for a new century by Steven Lewis Yaffee. Journal of Raptor Research 29:218-221.

Conservation Plans:

  • Block, W. M, F. Clemente, J. F. Cully, J. L. Dick, Jr., A. B. FRANKLIN, J. L. Ganey, F. P. Howe, W. H. Moir, S. L. Spangle, S. E. Rinkevich, D. L. Urban, R. Vahle, J. P. Ward, Jr., and G. C. White. 1995. Recovery plan for the Mexican Spotted Owl (Strix occidentalis lucida), Volume I. U. S. Fish and Wildlife Service, Albuquerque, New Mexico. 172pp.

Selected Technical Reports:

  • FRANKLIN, A. B. 2002. Comparison of occupancy probability estimates to habitat fitness potential in northern spotted owls in northwestern California. U. S. Fish and Wildlife Service, Arcata, California.
  • White, G. C., A. B. FRANKLIN, and D. C. Bowden. 2001. Research and development on new methods for monitoring Mexican spotted owl populations.  Rocky Mountain Research Station, Flagstaff, Arizona. 26pp.
  • White, G. C. and A. B. FRANKLIN. 2000. Estimation of Mexican spotted owl densities and population sizes in the Upper Gila Recovery Unit – prototype study. Rocky Mountain Research Station, Flagstaff, Arizona. 22pp.
  • FRANKLIN, A. B., K. P. Burnham, G. C. White, R. J. Anthony, E. D. Forsman, C. Schwarz, J. D. Nichols, and J. Hines. 1999. Range-wide status and trends in northern spotted owl populations. U. S. Geological Survey, Biological Resources Division, Corvallis, Oregon. 56pp. (reprint)
  • FRANKLIN, A. B., D. R. Anderson, and K. P. Burnham. 1998. Study of long-term trends in the annual probability of survival in North American birds. U. S. Geological Survey, Biological Resources Division Research Work Order 53, Denver, Colorado. 24pp.
  • FRANKLIN, A. B., K. R. Wilson, and T. M. Shenk. 1998. Estimated annual rate of change in a golden eagle population at the Altamont Pass Wind Resource Area, California. National Renewable Energy Laboratory, Golden, Colorado. 19pp.
  • Eberhardt, L. L., Nichols, J. D., Pollock, K. H, Noon, B. R, and A. B. FRANKLIN. 1993. A survey design to estimate the number of territorial spotted owls in Olympic National Park. Scientific Panel Review Final Report, Olympic National Park, Port Angeles, Washington. 29pp.
  • FRANKLIN, A.B. 1990. Analysis of timber industry density estimates for northern spotted owls in the inland Douglas-fir biome of northwest California. U.S. Fish and Wildlife Service, Portland, Oregon. 26pp.
  • FRANKLIN, A.B. 1989. A model illustrating territorial regulation in northern spotted owl populations. U. S. Fish and Wildlife Service, Portland, Oregon. 9pp.
  • FRANKLIN, A. B., and R. J. Gutiérrez. 1987. Population status of the northern spotted owl in northwest California: a demographic and empirical analysis. Northern. Spotted Owl Status Review Team, U.S. Fish and Wildlife Service, Portland, Oregon. 26pp.
  • Gutiérrez, R. J., J. P. Ward, A. B. FRANKLIN, and W. LaHaye. 1986. The influence of habitat heterogeneity on the dispersal pattern of juvenile spotted owls, Strix occidentalis caurina, in northwestern California. U.S. Forest Service, Portland, Oregon. 10pp.
  • Gutiérrez, R. J., and A. B. FRANKLIN. 1986. A review of wildlife planning in the draft land and resource management plan for the Shasta-Trinity National Forest. The Wilderness Society, Washington, DC, 14pp.

PRESENTATIONS

Conference Papers:

  • Shenk, T. M., D. Theobald, and A. B. Franklin. 2002. Movement patterns of lynx reintroduced to Colorado. The Wildlife Society 9th Annual Conference, Bismarck North Dakota.
  • Shenk, T. M., G. Byrne, G. C. White, A. B. FRANKLIN, and M. A. Wild. 2001. The Colorado lynx reintroduction program: an interim assessment. The Wildlife Society 8th Annual Conference, Reno, Nevada (poster presentation).
  • Shenk, T. M., G. C. White, and A. B. FRANKLIN. 2001. The Colorado lynx reintroduction program: an interim assessment. Colorado Wildlife Society 2001 Winter Conference, Grand Junction, Colorado.
  • FRANKLIN, A. B., K. P. Burnham, G. C. White, R. J. Anthony, E. D. Forsman, C. Schwarz, J. D. Nichols, and J. Hines. 2000. Trends in northern spotted owl populations: a meta-analytical approach. The Wildlife Society 7th Annual Conference, Nashville, Tennessee.
  • FRANKLIN, A. B., K. P. Burnham, and D. R. Anderson. 2000. Estimation of long-term trends and variation in avian survival probabilities using random effects models.  Euring 2000, Marshall, California.
  • FRANKLIN, A. B., K. P. Burnham, G. C. White, R. J. Anthony, E. D. Forsman, C. Schwarz, J. D. Nichols, J. Hines. 1999. Range-wide status and trends in northern spotted owl populations. 1999 Raptor Research Foundation Annual Meeting, La Paz, Mexico.
  • FRANKLIN, A. B. 1999. Exploring ecological relationships in survival and estimating rates of population change using program MARK. Second International Wildlife Management Congress, Budapest, Hungary.
  • FRANKLIN, A. B. and T. M. Shenk. 1999. The importance of parameter and variance component estimation in population viability analysis. Conference on Population Viability Analysis: Assessing Models for Recovering Endangered Species, San Diego, California.
  • White, G. C., and A. B. FRANKLIN. 1999. Use of marked animals for estimating parameters in PVA. Conference on Population Viability Analysis: Assessing Models for Recovering Endangered Species, San Diego, California.
  • White, G. C., J. P. Ward, Jr, R. J. Vahle, S. L. Spangle, S. E. Rinkevich, W. H. Moir, F. P. Howe, J. L. Ganey, A. B. FRANKLIN, J. L. Dick, and W. M. Block. 1999. Science versus reality in delisting criteria for an endangered species: The Mexican spotted owl experience. The 64th North American Wildlife and Natural Resources Conference, San Francisco, California.
  • Gutiérrez, R. J., J. P. Ward, A. B. FRANKLIN, P. Turchin, and W. S. Lahaye. 1998. Juvenile northern spotted owl dispersal: process and pattern. 1998 North American Ornithological Conference, St. Louis, Missouri.
  • Shenk, T. M., and A. B. FRANKLIN. 1996. Accuracy assessment of a vegetation map developed from Landsat Thematic Mapper imagery. 3rd Annual Conference of the Wildlife Society, Cincinnati, Ohio.
  • White, G. C., A. B. FRANKLIN, J. P. Ward, Jr., and W. M. Block. 1996. Monitoring population trends of Mexican Spotted Owls: necessary information for recovery of a threatened species. 114th Stated Meeting of the American Ornithologists' Union, Boise, Idaho.
  • FRANKLIN, A. B., D. R. Anderson, and R. J. Gutiérrez. 1995. Effects of climatic variation on survival rates in northern spotted owls. 80th Annual Meeting of the Ecological Society of America, Snowbird, Utah.
  • FRANKLIN, A. B., and T. M. Shenk. 1993. Meta-analysis as a tool for monitoring wildlife populations. International Wildlife Management Congress, San Jose, Costa Rica.
  • FRANKLIN, A. B. 1991. Population regulation in northern spotted owls: theoretical implications for management. Wildlife 2001: Populations, Oakland, California.
  • FRANKLIN, A. B., J. P. Ward, Jr., and R. J. Gutiérrez. 1987. Demography of the northern spotted owl in northwest California.. 105th Annual Meeting of American Ornithologists' Union, San Francisco, California.
  • Gutiérrez, R. J., J. P. Ward, A. B. FRANKLIN, and W. S. LaHaye. 1987. Dispersal ecology of juvenile northern spotted owls in northwestern California. 105th Annual Meeting of American Ornithologists' Union, San Francisco, California.
  • FRANKLIN, A. B. 1985. Breeding ecology of great gray owls in the Grand Teton region of Idaho and Wyoming. Raptor Research Foundation International Meeting on Management of Birds of Prey, Sacramento, California.
  • FRANKLIN, A. B. 1985. Breeding ecology of great gray owls in the Grand Teton region of Idaho and Wyoming. 54th Annual Meeting of Cooper Ornithological Society, Arcata, California.
  • Gutiérrez, R. J., J. P. Ward, Jr., A. B. FRANKLIN, and W. LaHaye. 1985. Dispersal ecology of juvenile northern spotted owls in northwest California. Raptor Research Foundation International Meeting on Management of Birds of Prey, Sacramento, California.
  • Gutiérrez, R. J., A. B. FRANKLIN, W. LaHaye, V. Meretsky, and J. P. Ward. 1984. Juvenile spotted owl dispersal in northwestern California: preliminary results. 54th Annual Meeting of the Cooper Ornithological Society, Humboldt State University, Arcata, California.
  • FRANKLIN, A. B., D. Clark, and D. Clark. 1978. Ecology and behavior of the Galapagos Rail. 96th Annual Meeting of American Ornithologists Union, Madison, Wisconsin.

Invited Seminars:

  • FRANKLIN, A. B. 2002. Forest fragmentation and habitat quality in spotted owls. Colloquium in the Life Sciences, Colorado State University, Fort Collins, Colorado.
  • FRANKLIN, A. B. 2002.  Science, politics, and spotted owls. Lida Scott Brown Ornithology Lecture, University of California - Los Angeles, Los Angeles, California.
  • FRANKLIN, A. B. 2002. Population dynamics of the California spotted owl: a meta-analysis. U. S. Forest Service, Sierra Nevada Forest Plan Amendment Review Team, Clovis, California
  • FRANKLIN, A. B. 2001. Climate, habitat quality, and population dynamics of northern spotted owls.  Department of Zoology and Physiology Seminar, University of Wyoming, Laramie, Wyoming.
  • FRANKLIN, A. B., D. R. Anderson, and K. P. Burnham. 2000. Estimation of long-term trends and variation in avian survival probabilities using random effects models.  Graduate-Faculty Seminar Series, Department Fishery and Wildlife Biology, Colorado State University, Fort Collins, Colorado.
  • FRANKLIN, A. B. 2000. Climate, habitat quality, and northern spotted owl populations: new avenues for research. Department of Wildlife and Fisheries Science, University of Arizona, Tucson, Arizona.
  • FRANKLIN, A. B. 2000. Climate, habitat quality, and population dynamics of northern spotted owls. Fisheries and Wildlife Seminar Series, University of Minnesota, St. Paul, Minnesota.
  • Carlson, P. C., A. B. FRANKLIN, R. J. Gutiérrez, and W. S. LaHaye. 2000. Divorce in spotted owls: why do they bother? Animal Ecology Series, Humboldt State University, Arcata, California.
  • FRANKLIN, A. B. 1999. Parameter estimation issues in demographic models. Sierra Nevada Framework Scientific Team, Sacramento, California.
  • FRANKLIN, A. B. 1999. Northern spotted owl demography in northwest California. Sierra Nevada Framework Scientific Team, Sacramento, California.
  • FRANKLIN, A. B. 1997. Factors affecting temporal and spatial variation in northern spotted owl populations. Department of Fisheries and Wildlife Biology Graduate-Faculty Seminar, Colorado State University, Fort Collins, Colorado.
  • FRANKLIN, A. B. 1989. Population biology of northern spotted owls in northwest California. Invited: Wildlife Seminar Series, Humboldt State University, Arcata, California.
  • FRANKLIN, A. B. 1989. Population parameters for northern spotted owls. Interagency Spotted Owl Management Committee, Portland, Oregon.
  • FRANKLIN, A. B. 1988. The spotted owl: an endangered species? Invited: Conservation Unlimited, Humboldt State University, Arcata, California.
  • FRANKLIN, A. B. 1988. The role of research in the spotted owl controversy. Invited: Moderated Panel on the Spotted Owl, Carter House Natural Science Museum, Redding, California.

Invited Symposia:

  • Block, W. M., A. B. FRANKLIN, J. L. Ganey, J. P. Ward, Jr., and G. C. White. 1999. Designing and conducting monitoring to evaluate success of restoration. 1999 Society for Ecological Restoration International Conference.
  • FRANKLIN, A. B. and R. J. Gutiérrez. 1999. Spotted Owls, forest fragmentation and forest heterogeneity. Symposium on Effects of Fragmentation on Western Bird Populations, 69th Annual Meeting of the Cooper Ornithological Society, Portland, Oregon.
  • Ganey, J. L., G. C. White, A. B. FRANKLIN, J. P. Ward, and W. M. Block. 1998. Monitoring population trend of Mexican spotted owls: necessary information for recovery of a threatened species. Mexican Spotted Owl Symposium, 1998 Raptor Research Foundation Annual Meeting, Ogden, Utah
  • FRANKLIN, A. B. 1998. What is habitat fragmentation? Symposium on Effects of Habitat Fragmentation on Wildlife, Colorado Chapter of The Wildlife Society 1998 Annual Meeting, Grand Junction, Colorado.
  • FRANKLIN, A. B., R. J. Gutièrrez, and P. C. Carlson. 1995. Trends in northern spotted owl populations on public lands in northwestern California. U. S. Fish and Wildlife Service Symposium on the Status and Management of the Northern Spotted Owl in California, Ukiah, California.
  • FRANKLIN, A. B. 1993. An overview of spotted owl studies and issues. WNAR Invited Papers 3 - Capture-recapture and northern spotted owls: statistics in action. 228th Meeting of the Institute of Mathematical Statistics, Laramie, Wyoming.
  • FRANKLIN, A. B., and E. Forsman. 1992. Future research directions in spotted owl population biology. Symposium on Demographics of the Northern Spotted Owl, 62nd Annual Meeting of Cooper Ornithological Society, Seattle, Washington.
  • FRANKLIN, A. B., J. P. Ward, Jr., R. J. Gutiérrez, and B. R. Noon. 1992. Demography of the northern spotted owl in northwest California. Symposium on Demographics of the Northern Spotted Owl, 62nd Annual Meeting of Cooper Ornithological Society, Seattle, Washington.
  • FRANKLIN, A. B., and J. P. Ward, Jr. 1992. Density of northern spotted owls: a meta-analysis. Spotted Owl Symposium, 1992 Annual Meeting of the Raptor Research Foundation, Bellevue, Washington.
  • FRANKLIN, A. B. and R. J. Gutiérrez. 1990. Demographics and management of the northern spotted owl. Theory and Tools for Managing Species at Risk, Univ. California, Davis, California.
  • FRANKLIN, A. B. 1987. Ecology and management of the northern spotted owl. 38th Annual Sierra-Cascade Logging Conference, Redding, California.

GRANTS AND CONTRACTS

Grants and contracts where listed as a Principal Investigator:

  • Gutiérrez, R. J. and A. B. FRANKLIN. 2001. Demography of the spotted owl in California. U. S. Forest Service, Region 5, San Francisco, California. $907,216
  • Gutiérrez, R. J. and A. B. FRANKLIN. 2000. Comparative demography of the spotted owl in California. U. S. Forest Service, Region 5, San Francisco, California. $224,923
  • White, G. C., A. B. FRANKLIN, and David Bowden. 2000. Research and Development on new approaches for monitoring Mexican spotted owl populations. Rocky Mountain Research Station, Flagstaff, Arizona. $41,273.
  • FRANKLIN, A. B. 2000. Technical assistance in developing northern spotted owl habitat map for California.  U. S. Fish and Wildlife Service, Arcata, CA. $4,960.
  • Anderson, D. R. and A. B. FRANKLIN. 1999. Population demography of northern spotted owls. Oregon Agricultural Research Foundation, Oregon State University, Corvallis, Oregon. $14,538
  • Gutiérrez, R. J. and A. B. FRANKLIN. 1999. Spotted owl demography studies - northwestern California. U. S. Forest Service, Region 5, San Francisco, California.  $233,987.
  • FRANKLIN, A. B. 1998.  Design and implementation of a mark-recapture study to monitor population trend in the Mexican spotted owl. U. S. Forest Service, Rocky Mountain Research Station, Flagstaff, Arizona. $9,566.
  • FRANKLIN, A. B. 1994. Analysis of Mexican spotted owl population data. U. S. Forest Service, Rocky Mountain Research Station, Flagstaff, Arizona   $2,499.
  • FRANKLIN, A. B. 1992. Review of Draft Conservation Strategy of the Mexican Spotted Owl. U. S. Forest Service, Rocky Mountain Research Station, Flagstaff, Arizona  $1,000
  • FRANKLIN, A. B. 1992. Development of Mexican spotted owl radio-telemetry study. U. S. Forest Service, Rocky Mountain Research Station, Flagstaff, Arizona. $2,000

Last updated: 1 October 2002